Fiche d'espèce de Copépode
Calanoida ( Ordre )
    Clausocalanoidea ( Superfamille )
        Scolecitrichidae ( Famille )
            Scolecitrichopsis ( Genre )
Scolecitrichopsis ctenopus  (Giesbrecht, 1888)   (F,M)
Syn.: Scolecithrix ctenopus Giesbrecht, 1888; 1892 (p.266, 285, 775, Descr.M, figs.M); T. Scott, 1894 b (p.48, figs.M); Giesbrecht & Schmeil, 1898 (p.46, Rem. M); Sars, 1925 (p.175); Farran, 1936 a (p.95, figs.F, Rem.); Bradford, 1973 (p.141, 146, Rem.); Campaner, 1979 (p.86, Rem.); Bradford & al., 1983 (p.103); Palomares Garcia & Vera, 1995 (tab.1); Kazmi, 2004 (p.229);
Scolecithricella spinipedata Mori, 1937 (1964) (p.53, Descr.F, figs.F); Chiba & al., 1955 (p.192); Chiba & al., 1957 (p.308); Delalo, 1968 (p.138);
Scolecithrix longicornis T. Scott, 1894 b (p.50, figs.F); Giesbrecht & Schmeil, 1898 (p.45); Sewell, 1948 (p.523);
Scolecithricella longicornis : A. Scott, 1909 (p.90, Rem.F);
Scolecithricella ctenopus : A. Scott, 1909 (p.91, Rem.M); Sewell, 1929 (p.212, figs.F); 1948 (p.384, 516, 523, 546, 557); Grice, 1962 (p.208, figs.F,M, juv., Rem.); Brodsky, 1962 c (p.125, figs.F,M); Tanaka, 1962 (p.53, figs.F,M); Chen & Zhang, 1965 (p.63, figs.M); Vervoort, 1965 (p.81, Rem.); Neto & Paiva, 1966 (p.24, Table III); Furuhashi, 1966 a (p.295, vertical distribution in Kuroshio region, Table 9); Owre & Foyo, 1967 (p.61, figs.F,M); Fleminger, 1967 a (tabl.1); Binet & al., 1972 (p.69); Bainbridge, 1972 (p.61, Appendix Table I: vertical distribution vs day/night, Table II: %); Carter, 1977 (1978) (p.35); Dessier, 1979 (p.205); Campaner, 1979 (p.86, Redef.); Sreekumaran Nair & al., 1981 (p.493), Stephen, 1984 (p.161, 169, Distribution vs thermocline & geographic); Guangshan & Honglin, 1984 (p.118, tab.); Brenning, 1984 (p.5, Rem.); 1985 a (p.28, Table 2); Longhurst, 1985 (tab.2); Madhupratap & Haridas, 1986 (p.105, tab.1); Jimenez-Perez & Lara-Lara, 1988; Hernandez-Trujillo, 1989 a (tab.1); Gopalakrishnan & Balachandran, 1992 (p.167, figs.1, 3, Table 1, 2); Shih & Young, 1995 (p.73); Padmavati & al., 1998 (p.349); Gopalakrishnan & Saraledevi, 1998 (p.134, fig.8, chart); Suarez-Morales & Gasca, 1998 a (p.111); Fernandez-Alamo & al., 2000 (p.1139, Appendix); Suarez-Morales & al., 2000 (p.751, tab.1); Madhupratap & al., 2001 (figs.4); Hernandez-Trujillo & Esqueda-Escargera, 2002 (in Appendix); Lopez-Ibarra & Palomares-Garcia, 2006 (p.63, Tabl. 1, seasonal abundance vs El-Niño); Jitlang & al., 2008 (p.65, Table 1); C.-Y. Lee & al., 2009 (p.151, Tab.2); Pillai H.U.K. & al., 2011 (p.239, Table 3, vertical distribution);
Scolecitrichopsus ctenopus : in CalCOFI regional list (MDO, Nov. 2013; M. Ohman, comm. pers.)
Vyshkvartzeva, 1999 (2000) (p.221); Ohtsuka & al., 2015 (p.123, Table 2: as S. stenopus).
Espèce Scolecitrichopsis ctenopus - Planche 1 de figures morphologiquesissued from : O. Tanaka in Publ. Seto Mar. Biol. Lab., 1962, X (1). [p.54, Fig.136]. As Scolecithricella ctenopus.
Female: a, habitus (left lateral side); b, forehead (dorsal); c, last thoracic segment and urosome (dorsal); d, P2; e, P5.

Male: f, 5th pair of legs.

Nota Female:
- Head and 1st pedigerous segment fused but line of fusion clearly xisible, 4th and 5th thoracic segments fused.
- Posterolateral margin of the last thoracic segment produced into a small spine.
- Cephalothorax abiut 4.6 rimes the abdomen length.
- Abdomen 4-segmented; segments and caudal rami in the proportional lengths 54 : 13 : 10 : 6 : 17 = 100.
- Genital segment about as long as wide; ventral surface swollen and sinuate.
- Caudal rami as long as wide
- A1 23-segmented; extends to the distal margin of the abdomen.
- A2 exopod slightly longer than endopod (5 : 4).
- Md with exopod about as long as endopod.
- Mx1 with 7 setae on the outer lobe; 5+2 setae on endopod; 3 setae on the 2nd basal.
- Mx2 with 3 long and 6 short vermiform filaments on the terminal segments; the spine ob the 5th lobe (= 5th endite) slender and long.
- P1 without outer edge spine on the 1st exopodal segment; basal segments with no small spines on the posterior surface; endopod with a number of scattered spines on the distal part of the segment in posterior aspect.
- P2 indented on the outer margin of 1st basal segment (= coxa); the outer edge spine on the 1st exopodal segment very small; terminal spine of exopod slightly distorted, with 15 teeth.
- P3 with a number of scattered small spines near the inner margin on the coxa; The outer margin deeply notched about the middle; basis with a row of small spines on the inner margin; Terminal spine of exopod with 15 teeth, the spine distorted.
- P5 with 2 free segments attached to the basal; 1st segment about as long as the 2nd; 2nd segment terminates into a strong spine, and with an inner marginal spine finely dentated on the external margin; the segment carries several small spines along the distal outer margin. the 1st segment furnished with small spines on the distal margin; segment with a row of minute spines on the distal 1/3 of the segment.

Nota Male:
- Abdomen contained 2.8 times in the length of the cephalothorax;
- Posterolateral corners of last thoracic segment not pointed.
- Line of fusion between head and 1st thoracic segment detectable on the middorsal region.
- A1 20-segmented on the right side; extends to the distal end of the abdomen.
- P5: right leg very small. Left leg very long and slender; endopod absent; distal segment with 2 apical spines of which the outer marginal one is long and curved; the inner proximal margin of the segment provided with a comb-like row of slender spines.

Espèce Scolecitrichopsis ctenopus - Planche 2 de figures morphologiquesissued from: Q.-c Chen & S.-z. Zhang in Studia Marina Sinica, 1965, 7. [Pl.20, 1-3]. As Scolecuthricella ctenopus.
Male (from E China Sea): 1, habitus (dorsal); 2, idem (lateral right side); 3, P5 (posterior).

Espèce Scolecitrichopsis ctenopus - Planche 3 de figures morphologiquesissued from : R.B.S. Sewell in Mem. Indian Mus., 1929, X. [p.214, Fig.79]. As Scolecithricella ctenopus.
Female (from S India): a, habitus (lateral left side); b, Mx2; c, Mxp; d, P1; e, P2; f, P3; f', idem (terminal spine enlarged); g, P5.

Espèce Scolecitrichopsis ctenopus - Planche 4 de figures morphologiquesissued from : T. Mori in The pelagic Copepoda from the neighbouring waters of Japan, 1937 (2nd edit., 1964). [Pl.26, Figs.11-16]. As Scolecithricella spinipedata.
Female: 11, P2 (posterior); 12, habitus (lateral); 13, P5; 14, genital area; 15, P3 (posterior); 16, P4 (posterior).

Espèce Scolecitrichopsis ctenopus - Planche 5 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. - Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf. 13, Figs;36, 37, 38 ]. As .
Male: 36, P5 (posterior view); 37, right P5; 38, exopodite 2 and 3 of left leg of P5.

Espèce Scolecitrichopsis ctenopus - Planche 6 de figures morphologiquesIssued from : W. Giesbrecht in Systematik und Faunistik der Pelagischen Copepoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. – Fauna Flora Golf. Neapel, 1892. Atlas von 54 Tafeln. [Taf.37, Fig15]. As Scolecithrix ctenopus.
Male: habitus (lateral).

Espèce Scolecitrichopsis ctenopus - Planche 7 de figures morphologiquesissued from : G.D. Grice in Fish. Bull. Fish and Wildl. Ser., 1962, 61. [p.207, Pl.16, Figs.6-15]. As Scolecithricella ctenopus.
Female juvenile (from equatorial Pacific): 6, habitus (dorsal); 7, P5.

Female adult: 8-9, habitus (dorsal and lateral, respectively); 10, posterior part of thorax and urosome (dorsal); 11, same (left side); 12, same (right side); 13, rostrum; 14, P5.
Nota: The female specimens fall into two size groups, the smaller of which is very similar to Farrn's small specimen. The head and thorax in this form are separated by a fine line (in Farran's specimen they were fused) and a faint line is also seen to separate the 4th and 5th thoracic segments (fig.6). Althought the P5 are fairly well diifferentiated, the genital segment is undevelopped in this small specimen. In the larger form (figs.8-14) the head and 1st thoracic segment are incompletely fused and the 4th and 5th thoracic segments are completely fused.

Male: 15, P5.

Espèce Scolecitrichopsis ctenopus - Planche 8 de figures morphologiquesissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.60, Fig;378]. As Scolecithricella ctenopus.
Female14°55'-15°00'N, 64°50'-67°05'W): 378, habitus (lateral).

Espèce Scolecitrichopsis ctenopus - Planche 9 de figures morphologiquesissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.25, Fig.113]. As Scolecithricella ctenopus.
Female: P5.

Espèce Scolecitrichopsis ctenopus - Planche 10 de figures morphologiquesissued from : H.B. Owre & M. Foyo in Fauna Caribaea, 1, Crustacea, 1: Copepoda. Copepods of the Florida Current. [p.23, Figs.95-97]. As .
Male: P5; 96, detail of P5; 97, distal segment of left P5.

Ref. compl.:
Uysal & al., 2002 (p.18, tab.1); Kuriyama & Nishida, 2006 (p.300: Tab.II); Fernandes, 2008 (p.465, Tabl.2); Cornils & al., 2010 (p.2076, Table 3); Schnack-Schiel & al., 2010 (p.2064, Table 2: E Atlantic subtropical/tropical); Dorgham & al., 2012 (p.473, Table 4: abundance vs season);
NZ: 15

Carte de distribution de Scolecitrichopsis ctenopus par zones géographiques
Espèce Scolecitrichopsis ctenopus - Carte de distribution 3issued from : Gopalakrishnan & Saraledevi in Pelagic Biogeography ICoPB II. Proceedings of the 2nd International Conference. Final report of SCOR/IOC working group 93. Noordwijkerhout, The Netherlands 9-14 July 1995. Workshop Report No.142. UNESCO, 1998. [p.134, Fig.8].
Distribution and abundance in number of specimens/standard haul of Scolecithricella ctenopus (= Scolecitrichopsis ctenopus) in the Indian Ocean.
Espèce Scolecitrichopsis ctenopus - Carte de distribution 4issued from : U. Brenning in Wiss. Z. Wilhelm-Pieck-Univ. Rostock - 33. Jahrgang 1984. Mat.-nat. wiss. Reihe, 6. [p.6, Fig.2].
Spatial distribution for Scolecithricella ctenopus (= Scolecithrix ctenopus) and other scolecithrids from 8° S - 26° N; 16°- 20° W, for different expeditions (V1: Dec. 1972- Jan. 1973; V2: Feb/Mar. 1973; VI: May 1974; IV: Jun./Jul. 1972).
South Africa (E), Angola (Baia Farta), Congo, G. of Guinea, Ivorian shelf, off Senegal, off Amazon, Venezuela, Caribbean Sea, Caribbean, Florida, E Medit. (Lebanon Basin), Sharm El-Sheikh, Red Sea, Arabian Sea, off S Laccadive Is., Natal, Indian (N, SW & equatorial, E India, Bay of Bengal, Andaman Sea (Batten Island), Indonesia-Malaysia, SW Celebes; China Seas (East China Sea, South China Sea), Taiwan (NW), Japan, Bikini Is., Pacif. (W equatorial), Australia (Great Barrier), Pacif. (equatorial), California, W Baja California, Bahia Magdalena (rare), G. of California, W Mexico, G. of Tehuantepec, off W Guatemala
N: 64
(16) F: 1,49-1,43; M: 1,65-1,45; (29) F: 1,26; (34) F: 1,47; 1,25; (47) M: 1,3; (57) F: 1,54; M: 1,83; (101) F: 1,53-1,46; M: 1,54; 1,46; (117) F: 1,52; M: 1,57-1,52; (150) F: 1,65-1,52; M: 1,65-1,6; (290) M: 1,9; (1000) F: 1,6; M: 1,6; (1111) M; 1,68; {F: 1,25-1,65; M: 1,30-1,90}
Rem.: épipélagic. warm-water inhabitant.
After Tanaka (1962, p.55)Scolecithricella spinipedata Mori, 1937 appears to be identical with S. ctenopushaving a small outer edge spine on the 1st exopodal segment of P2. The female P5 shows some variations in the length of the inner marginal spine.For Tanaka (1962, p.55) Scolecithricella ctenopus appears to be intermediate between Scolecithricella and Amallothrix in having only vermiform appendages on Mx2, and a cylindrical P5 in the female. The male P5 differs also greatly from that found in the male of Scolecithricella stricto sensu.
Dernière mise à jour : 19/03/2017

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Razouls C., de Bovée F., Kouwenberg J. et Desreumaux N., 2005-2019. - Diversité et répartition géographique chez les Copépodes planctoniques marins. Sorbonne Université, CNRS. Disponible sur [Accédé le 22 janvier 2019]

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